Archaeocetes:
Archaic Whales, Dolphins and Porpoises

Last Updated 06-23-04


Important because they :
- Represent the earliest radiation of cetaceans
- Include ancestors of two modern suborders of cetaceans (Mysticeti and Odontoceti)
- Document the initial amphibious stages in cetacean evolution

Six families of cetaceans are commonly included in Archaeocetes.
- Pakicetidae
- Ambulocetidae
- Remingtonocetidae
- Protocetidae
- Basilosauridae
- Dorudontidae

These archaeocete families document that the Eocene cetacean evolution is characterized by increasing aquatic adaptations, starting at amphibious early whales (pakicetids, ambulocetids) to more marine protocetids.

- Characters of modern and fossil whales
- heavy ear ossicles (malleus, incus, stapes)
- bone forming the wall of ear (tympanic) with thick internal lip (involucrum) brand wavy crest (sigmoid process) externally
- Simplified cheek teeth
. Characters of modern whales (may or may not be present in fossils)
- Tail fluke
- Forelimb modified to a flipper
- Streamlined body design
- No hair as adults


Pakicetidae

A reconstruction of Pakicetus.
Illustration by Carl Buell, and taken from http://www.neoucom.edu/Depts/Anat/Pakicetid.html

- Early-to-middle Eocene Period, approximately 50 million years ago in India and Pakistan
- Dozens of fossils known, but none consist of complete skeletons
- Known skeletal elements include skulls, teeth, and jaw fragments
- Fossils only found in freshwater deposits, most represent shallow ephemeral streams arid climates
- Although pakicetids were land mammals, it is clear that they are related to whales and dolphins based on a number of specializations of the ear.
- Genera included:
- Pakicetus
P. inachus - 1981
P. attocki - 1980
- Nalacetus
N. ratimitus - 1980
- Ichthyolestes
I. pinfoldi - 1958


Ambulocetidae

The skeleton of Ambulocetus natans (approximately 12 feet long).
Illustration taken from http://www.neoucom.edu/Depts/Anat/Pakicetid.html

- Known from middle Eocene Period (49 million years ago) rocks in northern India and Pakistan
- Fewer than 10 described fossils; one consists of a nearly complete skeleton (Ambulocetus natans)
- A. natans is the primary source of information about early cetacean biology
- A. natans skeleton suggest it was probably slow on land and an ambush hunter which attacks prey in or near shallow water (like modern crocodiles)
- Only known from nearshore marine environments, including bays and estuaries
- Genera included:
- Ambulocetus
A. natans - 1994
- Gandakasia
G. potens - 1958
- Himalayacetus
H. subathuensis - 1998


Remingtonocetidae

Reconstruction of the remingtonocetid Kutchicetus.
It was a small animal, no bigger than a river otter.
Illustration by Carl Buell and taken from http://www.neoucom.edu/Depts/Anat/Remi.html.

- Only known from India and Pakistan sediments approximately 46 to 43 million years ago (middle Eocene Period)
- Dozens of fossils described, most document morphology of skull and lower jaw
- Oldest and most primitive fossil (Attockicetus) found with Ambulocetus
- All other fossils known from marine, nearshore deposits; may have lived in bays and saltwater swamps
- Genera included:
- Andrewsiphius
A. sloani - 1972
A. kutchensis - 1975 (subsumed to A. sloani - 2001)
A. minor - 1975 (subsumed to A. sloani - 2001)
- Remingtonocetus
R. harudiensis - 1975
R. domandaensis - 2001
- Dalanistes
D. ahmedi - 1995
- Attockicetus
A. praecursor - 2000
- Kutchicetus
K. minimus - 2000 (subsumed to Andrewsiphius sloani - 2001)


Protocetidae

Partial skeleton of Rodhocetus.

- Found in middle Eocene rocks in Indo-Pakistan, Africa, Europe, and North America
- Oldest whales to disperse across the oceans; probably only warm seas near tropics
- Several partial skeletons described
- Indo-Pakistan protocetids inhabited same environments as the remingtonocetids
- Protocetids from other continents known from shallow marine environments
- First whale group to develop tail flukes and most were probably agile, quick aquatic hunters who preyed on small animals
- Genera included:
Subfamily Protocetinae
- Protocetus
P. atavus - 1904
- Eocetus
E. wardii - 1999
E. schweinfurthi - 1904
- Pappocetus
P. lugardi - 1920
- Babiacetus
B. indicus - 1984
B. mishrai - 1998
- Takracetus
T. simus - 1995
- Georgiacetus
G. vogtlensis - 1998
- Natchitochia
N. jonesi - 1998
Subfamily Indocetinae
- Indocetus
I. ramani - 1975
- Rodhocetus
R. kasrani - 1994
R. balochistanensis
-Qaisracetus
Q. arifi - 2001
-Artiocetus
A. clavis


Basilosauridae

Reconstruction of Basilosaurus cetoides .
Illustration taken from http://darla.neoucom.edu/DEPTS/ANAT/BasilAndDor.htm.

Dorudontidae

Reconstruction of Zygorhiza kochii (not to scale).
Illustration taken from http://darla.neoucom.edu/DEPTS/ANAT/BasilAndDor.htm.

Basilosaurids and dorudontids lived in the late Eocene, approximately 35 and 41 million years ago. They are mainly known from the eastern United States and from Egypt, but were probably worldwide in their distribution. Basilosaurids were enormous (possibly up to 60 feet long) and had snake-like bodies. They had a tail fluke. Dorudontids were proportionally more like dolphins. Both basilosaurids and dorudontids had complete hind limbs that included a mobile knee and several toes. However these extremities were tiny, so small that they were certainly not important in aquatic propulsion.
- Genera included:
Subfamily Basilosauriniae
- Basilosaurus
B. cetoides - 1839 (synonym Zeuglodon cetoides)
B. isis - 1904
B. hussaini - 1997 (or Basiloterus hussaini)
- Basiloterus
B. drazindai - 1997 (or Basilosaurus drazindai)
- Gaviacetus
G. razai - 1995
G. sahnii - 1998
Subfamily Dorudontinae
- Dorudon
D. atrox - 1906 (formerly Prozeuglodon atrox)
D. serratus - 1845
- Zygorhiza
Z. kochii - 1847
Z. species . 1997
- Saghacetus
S. osiris - 1894
(formerly Dorudon osiris, D. zitteli, D. sensitivius, D. elliotsmithii)
- Ancalecetus
A. simonsi - 1996
- Chrysocetus
C. healyorum - 2001
- Pontogeneus
P. priscus
P. brachyspondylus (formerly Zeuglodon brachyspondylus)


|Home| |Officers| |Calendar| |WCRC Constitution|
|WCRC helps ARK| |WCRC helps BREC| |WCRC Yahoo Groups| |Links|
|GOM Sea Turtles| |Sea Turtle Comparison| |Sea Turtle Morphology| |Sea Turtle Glossary|
|GOM Cetaceans| |World Cetaceans| | Archaic Cetaceans|
|Cetacean Anatomy| |Cetacean Glossary|

Source of information: Thewissen, J.G.M. "Hans", Ph.D. http://darla.neoucom.edu/DEPTS/HRES/FACULTY/BMSfac/ThewissenJ.html. 11/03
http://archaeocete.org/. 06/04

e-mail questions, comments, and/or suggestions to
wcrc@sci.tamucc.edu

Webmaster: Candice "Orca" Mottet

This page was created by: Candice "Orca" Mottet